Johnson, K., and Clyde, W., 2016, Ancient Wyoming: A Dozen Lost Worlds Based on the Geology of the Bighorn Basin: Fulcrum Publishing, Golden, CO, 64p, ISBN-13: 978-1936218172.

 Journal Papers

Fuentes, A.J., Clyde, W.C., Weissenburger, K., Bercovici, A., Lyson, T.R., Miller, I.M., Ramezani, J., Schmitz, M.D., and Johnson, K.R., 2019, Constructing a timescale of biotic recovery across the Cretaceous–Paleogene boundary, Corral Bluffs, Denver Basin, Colorado: Rocky Mountain Geology   54 (2): 133–153, 10.24872/rmgjournal.54.2.133. (PDF)

Lyson, T.R., Miller, I.M., Bercovici A.D., Weissenburger1, K., Fuentes, A.J., Clyde, W.C., Hagadorn, J.W., Butrim, M.J., Johnson, K.R., Fleming, R.F., Barclay, R.S., Maccracken, S.A., Lloyd, B., Wilson, G.P., Krause, D.W., Chester., S.G.B., 2019, An exceptional continental record of biotic recovery after the Cretaceous–Paleogene mass extinction, Science 366: 977–983, 10.1126/science.aay2268. (PDF)

Currano, E.D., Pinheiro, E.R.S., Buchwaldt, R., Clyde, W.C., and Miller, I.M., 2019, Endemism in Wyoming plant and insect herbivore communities during the early Eocene hothouse: Paleobiology, p. 1–19, doi:10.1017/pab.2019.18. (PDF)

Murphey, P. C., Kelly, T. S., Chamberlain, K. R., Tsukui, K., and Clyde, W. C. 2018. Mammals from the earliest Uintan (middle Eocene) Turtle Bluff Member, Bridger Formation, southwestern Wyoming, USA, Part 3: Marsupialia and a reevaluation of the Bridgerian-Uintan North American Land Mammal Age transition. Palaeontologia Electronica 21.2.24A 1-52. https: // (Link)

Westerhold, T., Röhl, U., Wilkens, R.H., Gingerich, P.D., Clyde, W.C., Wing, S.L., Bowen, G.J., and Kraus, M.J., 2018, Synchronizing early Eocene deep-sea and continental records – cyclostratigraphic age models for the Bighorn Basin Coring Project drill cores, Climate of the Past, v. 14, p. 303–319, doi: 10.5194/cp-14-303-2018. (Link)

Widlansky, S.J., Clyde, W.C., O’Connor, P.M., Roberts, E.M., and Stevens, N.J., 2018, Paleomagnetism of the Cretaceous Galula Formation and implications for vertebrate evolution, Journal of African Earth Sciences, v. 139, p. 403–420, doi: 10.1016/j.jafrearsci.2017.11.029. (PDF)

Phillips, S. C., Johnson, J. E., Clyde, W. C., Setera, J. B., Maxbauer, D. P., Severmann, S., and Riedinger, N., 2017, Rock magnetic and geochemical evidence for authigenic magnetite formation via iron reduction in coal-bearing sediments offshore Shimokita Peninsula, Japan (IODP Site C0020), Geochemistry, Geophysics, Geosystems 18, 2076–2098, doi:10.1002/2017GC006943. (PDF)

D'Ambrosia, A. R., Clyde, W. C., Fricke, H. C., Gingerich, P. D., Abels, H. A., 2017, Repetitive mammalian dwarfing during ancient greenhouse warming events, Science Advances, 3 (3), e1601430, doi: 10.1126/sciadv.1601430. (PDF)

Krause, J. M., Clyde, W. C., Ibañez-Mejía, M. Schmitz, M. D., Barnum, T., Bellosi, E. S., Wilf, P., 2017, New age constraints for early Paleogene strata of central Patagonia, Argentina: implications for the timing of South America land mammal ages, Geological Society of America Bulletin, 129(7-8):886-903, doi: 10.1130/B31561.1. (PDF)

Clyde, W. C., Ramezani, J., Johnson, K. R., Bowring, S. A., Jones, M. M. 2016, Direct high-precision U-Pb geochronology of the end-Cretaceous extinction and calibration of Paleocene astronomical timescales. Earth and Planetary Science Letters, 452:272-280, doi: 10.1016/j.epsl.2016.07.041. (PDF)

Maxbauer, D. P., Feinberg, J. M., Fox, D. L., Clyde, W. C., 2016, Magnetic minerals as recorders of weathering, diagenesis, and paleoclimate: a core-outcrop comparison of Paleocene-Eocene paleosols in the Bighorn Basin, WY, U.S.A. Earth and Planetary Science Letters 452:15-26, doi: 10.1016/j.epsl.2016.07.029. (PDF)

Comer, E., Slingerland, R., Krause, J., Iglesias. A., Clyde, W. C., Wilf, P., and Raigemborn, M., 2015, Sedimentary facies and depositional environments of diverse early Paleocene floras in the north-central San Jorge Basin (Patagonia, Argentina), Palaios 30, 553–573, DOI: doi: 10.2110/palo.2014.064. (PDF)

Machlus, M.L., Ramezani, J., Bowring, S.A., Hemming, S.R., Tsukui, K., and Clyde, W.C., 2015, A strategy for cross-calibrating U–Pb chronology and astrochronology of sedimentary sequences: An example from the Green River Formation, Wyoming, USA: Earth and Planetary Science Letters 413: 70–78, doi: 10.1016/j.epsl.2014.12.009. (PDF)

Bowen, G.J., Maibauer, B.J., Kraus, M.J., Rohl, U., Westerhold, T., Steimke, A., Gingerich, P.D., Wing, S.L., and Clyde, W.C., 2015, Two massive, rapid releases of carbon during the onset of the Palaeocene-Eocene thermal maximum: Nature Geoscience 8: 44–47; doi:10.1038/ngeo2316. (PDF)

D’Ambrosia, A.R., Clyde, W.C., Fricke, H.C., and Chew, A.E., 2014, Stable isotope patterns found in early Eocene equid tooth rows of North America: Implications for reproductive behavior and paleoclimate: Palaeogeography, Palaeoclimatology, Palaeoecology 414:310–319; doi: 10.1016/j.palaeo.2014.09.014. (PDF)

Clyde, W. C., Wilf, P., Iglesias, A., Slingerland, R. L., Barnum, T., Bijl, P. K., Bralower, T. J., Brinkhuis, H., Comer, E. E., Huber, B. T., Ibañez-Mejia, M., Jicha, B. R., Krause, J. M., Schueth, J. D., Singer, B. S., Raigemborn, M. S., Schmitz, M. D., Sluijs, A., Zamaloa, M. del C., 2014, New age constraints for the Salamanca Formation and lower Río Chico Group in the western San Jorge Basin, Patagonia, Argentina: Implications for K/Pg extinction recovery and land mammal age correlations, Geological Society of America Bulletin 126: 289–306; doi:10.1130/B30915.1. (PDF)

Clyde, W. C., Gingerich, P. D., Wing, S. L., Röhl, U., Westerhold, T., Bowen, G., Johnson, K., Baczynski, A. A., Diefendorf, A., McInerney, F., Schnurrenberger, D., Noren, A., Brady, K., and the BBCP Science Team. 2013. Bighorn Basin Coring Project (BBCP): a continental perspective on early Paleogene hyperthermals. Scientific Drilling 16:21-31; doi:10.5194/sd-16-21-2013. (PDF)

Khan, I.H. and Clyde, W.C. 2013. Lower Paleogene tectonostratigraphy of Balochistan: Evidence for time-transgressive Late Paleocene-Early Eocene uplift. Geosciences, 3, 466-501 ; doi:10.3390/geosciences3030466. (PDF)

Abels, H. A., Clyde, W. C., Gingerich, P. D., Hilgen, F. J., Fricke, H. C., Bowen, G. J. and Lourens, L. J. 2012. Terrestrial carbon isotope excursions and biotic change during Palaeogene hyperthermals. Nature Geoscience 5:326-329; doi: 10.1038/NGEO1427. (PDF)

Tsukui, K., Clyde, W. C. 2012. Fine-tuning the calibration of the early to middle Eocene geomagnetic polarity time scale: Paleomagnetism of radioisotopically dated tuffs from Laramide foreland basins. Geological Society of America Bulletin 124:870-885; doi: 10.1130/B30545.1. (PDF)

Clyde, W. C., Wing, S. L., Gingerich, P. D. 2012. Coring Project in Bighorn Basin: Drilling Phase Complete, Eos: Transactions of the American Geophysical Union, 93(4), 41-42. (PDF)

Roberts, E. M., Stevens, N. J., O’Connor, P. M., Dirks, P. H. G. M., Gottfried, M. D., Clyde, W. C., Armstrong, R. A., Kemp, A. I. S., and Hemming, S. 2012. Initiation of the western branch of the East African Rift coeval with the eastern branch. Nature Geoscience 5:289-294; doi: 10.1038/NGEO1432. (PDF)

Ting, S., Wang, Y., Clyde, W. C., Koch, P. L., Tong Y., Meng J., Bowen, G., Li, Q., and Snell, K. E. 2011. Asian early Paleogene chronology and mammalian faunal turnover events. Vertebrata PalAsiatica 49: 1-28 (PDF)

Clyde W. C., Ting S., Snell K. E., Bowen, G. J., Tong, Y., Koch, P. L., Li, Q., Wang, Y. 2010. New Paleomagnetic and Stable-Isotope Results from the Nanxiong Basin, China: Implications for the K/T Boundary and the Timing of Paleocene Mammalian Turnover. Journal of Geology 118: 131-143. (PDF)

Clyde, W. C., Tong, Y., Snell, K. E., Bowen, G. J., Ting, S., Koch, P. L., Li, Q., Wang, Y., Meng, J. 2008. An integrated stratigraphic record from the Paleocene of the Chijiang Basin, Jiangxi Province (China): Implications for mammalian turnover and Asian block rotations. Earth and Planetary Science Letters 269: 553-563. (PDF)

Gunnell, G. F., Gingerich, P. D., Ul-Haq, M., Bloch, J. I., Khan, I. H., Clyde, W. C. 2008. New Primates (Mammalia) from the early and middle Eocene of Pakistan and their paleobiogeographical implications, Contributions from the Museum of Paleontology, University of Michigan 32: 1-14. (PDF)

Clyde, W. C., Hamzi, W., Finarelli, J. A., Wing, S. L., Schankler, D., Chew, A. 2007. Basin‑wide magetostratigraphic framework for the Bighorn Basin, Wyoming. Geological Society of America Bulletin 119: 848B859. (PDF)

Ting, S., Meng, J., Qian, L. Wang, Y., Tong, Y., Shiebout, J. A., Koch, P.L., Clyde, W.C., Bowen, G. J.2007. Ganungulatum xincunliense, an artiodactyl-like mammal (Ungulata, Mammalia) from the Paleocene,Chijiang Basin, Jiangxi, China.  Vertebrata PalAsiatica 45:279-286. (PDF)

Secord, R., Gingerich, P. D., Smith, M. E., Clyde, W. C., Wilf, P., and Singer, B. S. 2006. Geochronology and mammalian biostratrigraphy of middle and upper Paleocene continental strata, Bighorn Basin, Wyming. American Journal of Science 306, 211‑245. (PDF)

Seiffert, E. R., Simons E. L., Clyde W. C., Rossie J. B., Attia, Y., Bown T. M., Chatrath, P., Mathison M. E. 2005. Basal Anthropoids from Egypt and the Antiquity of Africa=s Higher Primate Radiation. Science 310: 300‑304. (PDF)

Clyde, W. C., Finarelli, J. A., Christensen, K. E. 2005. Evaluating the relationship between pedofacies and faunal composition; Implications for faunal turnover at the Paleocene‑Eocene boundary. Palaios 20: 390‑399. (PDF)

Finarelli, J. A. and Clyde, W. C.  2004.  Reassessing hominoid phylogeny: Evaluating congruence in the morphologic and temporal data.  Paleobiology 30: 614‑651. (PDF)

Clyde, W. C., Bartels, W. S., Gunnell, G. F., Zonneveld, J.-P.  2004.  Discussion: 40Ar/39Ar geochronology of the Eocene Green River Formation, Wyoming.  Geological Society of America Bulletin 116:251-252. (PDF)

Ting, S., Wang, Y., Schiebout J. A., Koch, P., Clyde, W. C., Bowen, G. J., Wang, Y. 2004. New early Eocene mammalian fossils from the Hengyang Basin, Hunan, China.  Bulletin of Carnegie Museum of Natural History 36, p. 291‑301. (PDF)

Clyde, W. C., Khan, I. H., Gingerich, P. D. 2003.  Stratigraphic response and mammalian dispersal during initial India-Asia collision: Evidence from the Ghazij Formation, Baluchistan, Pakistan. Geology 31:1097-1100. (PDF)

Clyde, W. C. and Christensen, K.  2003.  Testing the relationship between pedofacies and avulsion using markov analysis.  American Journal of Science 303:60-71. (PDF)

Bowen, G. J., Clyde, W. C., Koch, P. L., Ting, S., Alroy, J., Takehisa, T., Wang, Y., Wang, Y. 2002.  Mammalian dispersal at the Paleocene/Eocene boundary.  Science 295:2062-2065. (PDF)

Finarelli, J. A. and Clyde, W. C.  2002.  Comparing the gap excess ratio and the retention index of the stratigraphic character.  Systematic Biology 51:166-176. (PDF)

Clyde, W. C., Sheldon, N. D., Koch, P. L., Gunnell, G. F., and Bartels, W. S.  2001.  Linking the Wasatchian-Bridgerian boundary to the Global Cenozoic Climate Optimum: new magnetostratigraphic and isotopic results from South Pass, Wyoming.  Palaeogeography, Palaeoclimatology, Palaeoecology 167:175-199. (PDF)


Gingerich, P. D., Arif, M., Khan, I. H., Clyde, W. C., and Bloch, J. I.  1999.  Machocyon abbasi, a new early Eocene Quettacyonid (Mammalia, Condylarthra) from the middle Ghazij Formation of Mach and Deghari Coal fields, Baluchistan (Pakistan).  Contributions from the Museum of Paleontology, University of Michigan 30:233-250. (PDF)

Zaprowski, B. J., and Clyde, W. C.  1999.  Playing Wegener in a mock world: A laboratory exercise for introductory Earth Science classes.  Journal of Geoscience Education 47:336-340. (PDF)


Fricke, H. C., Clyde, W. C., and O'Neil, J. R.  1998.  Intra-tooth variations in δ18O(PO4) of mammalian tooth enamel as a record of seasonal variations in continental climate variables.  Geochimica et Cosmochimica Acta 62:1839-1850. (PDF)

Clyde, W.C., and Gingerich, P. D.  1998.  Mammalian community response to the latest Paleocene thermal maximum: An isotaphonomic study from the northern Bighorn basin, Wyoming.  Geology 26:1011-1014. (PDF)

Fricke, H. C., Clyde, W. C., O'Neil, J. R., and Gingerich, P. D.  1998.  Evidence for rapid climate change in North America during the latest Paleocene thermal maximum: oxygen isotope compositions of biogenic phosphate from the Bighorn Basin (Wyoming), Earth and Planetary Science Letters 160:193-208. (PDF)

Clyde, W. C., Zonneveld, J-P., Stamatakos, J., Gunnell, G. F., and Bartels, W. S.  1997.  Magnetostratigraphy across the Wasatchian-Bridgerian boundary in the Green River Basin, Wyoming.   Journal of Geology 105:657-669. (PDF)

Clyde, W. C., and Fisher, D. C.  1997.  Comparing the fit of stratigraphic and morphologic data in phylogenetic analysis.  Paleobiology 23:1-19. (PDF)

Clyde, W. C., Stamatakos, J., and Gingerich, P. D.  1995.  Chronology of the Wasatchian land-mammal age: magnetostratigraphic results from the McCullough Peaks section, northern Bighorn Basin, Wyoming: A reply.  Journal of Geology 103:464-466. (PDF)

Gingerich, P. D., Arif, M., and Clyde, W. C.  1995.  New Archaeocetes (Mammalia, Cetacea) from the middle Eocene Domanda Formation of the Sulaiman Range, Punjab (Pakistan).  Contributions from the Museum of Paleontology, University of Michigan 29:291-330. (PDF)

Clyde, W. C., Stamatakos, J., and Gingerich, P. D.  1994.  Chronology of the Wasatchian land-mammal age: magnetostratigraphic results from the McCullough Peaks section, northern Bighorn Basin, Wyoming.  Journal of Geology 102:367-377. (PDF)

Clyde, W. C., and Gingerich, P. D.  1994.  Rates of evolution in the dentition of early Eocene Cantius: comparison of size and shape.  Paleobiology 20:506-522. (PDF)

Book and Monograph Chapters

LeCain, R., Clyde, W.C., Wilson, G.P., and Riedel, J., 2014, Magnetostratigraphy of the Hell Creek and lower Fort Union Formations in northeastern Montana, in Wilson, G.P., Clemens, W.A., Horner, J.R., and Hartman, J.H., eds., Through the End of the Cretaceous in the Type Locality of the Hell Creek Formation in Montana and Adjacent Areas: Geological Society of America Special Paper 503, p. 137–147, doi:10.1130/2014.2503(04). (PDF)

Clyde, W. C., LeCain, R., 2012, Terrestrial Ecosystem Response to Climate Change during the Paleogene in Hannah, L. (ed.) Saving a Million Species: Extinction Risk from Climate Change, Island Press, pp. 157-177. (PDF)

Seiffert E.R., Bown T.M., Clyde W.C., and Simons E.L. 2008. Geology, paleoenvironment, and age of Birket Qarun Locality 2 (BQ-2), Fayum Depression, Egypt in Fleagle, J.G and Gilbert, C. C. (eds) Elwyn L. Simons: A Search for Origins. New York: Springer, pp. 71-86. (PDF)

Robinson, P., Gunnell, G. F., Walsh, S. L., Clyde, W. C., Storer, J. E., Stucky, R. K., Froehlich, D. J., Ferrusquia-Villafranca, I., .McKenna, M. C. (2004). Wasatchian through Duchesnean biochronology in Woodburne, M. O. (ed.), Late Cretaceous and Cenozoic Mammals of North America: Biostratigraphy and Geochronology, Columbia University Press, New York, pp. 106-155. (PDF)

Zonneveld, J‑P., Bartels, W.S. and Clyde, W.C.  2003.  Stratal architecture of an Early Eocene fluvial‑lacustrine depositional system, Little Muddy Creek area, southwestern Green River Basin, Wyoming.  The Cenozoic of the Rocky Mountain Region, RMS‑SEPM Special Publication: 253‑288. (PDF)

Koch, P. L, Clyde, W. C, Hepple, R. P. Fogel, M. L., Wing, S. L., Zachos, J. C.  2003. Carbon and oxygen isotope records from paleosols spanning the Paleocene-Eocene boundary, Bighorn Basin, Wyoming in Wing, S.L., Gingerich, P.D., Schmitz, B., and Thomas, E. (eds.) Causes and Consequences of Globally Warm Climates in the Early Paleogene, Geological Society of America, Special Paper 369, p.49-64. (PDF)

Ting, S., Bowen, G. J., Koch, P. L., Clyde, W. C., Wang, Y., Wang, Y., McKenna, M. C. 2003. Biostratigraphic, chemostratigraphic, and magnetostratigraphic study across the Paleocene‑Eocene boundary in the Hengyang Basin, Hunan, China. in Wing, S.L., Gingerich, P.D., Schmitz, B., and Thomas, E. (eds.) Causes and Consequences of Globally Warm Climates in the Early Paleogene, Geological Society of America, Special Paper 369, p.521-535. (PDF)


Gingerich, P. D. and Clyde, W. C.  2001.  Overview of mammalian biostratigraphy in the Paleocene-Eocene Fort Union and Willwood Formations of the Bighorn and Clarks Fork Basins.  In Gingerich (ed.) Paleocene-Eocene Stratigraphy and Biotic Change in the Bighorn and Clarks Fork Basins, Wyoming, University of Michigan Papers on Paleontology 33: 1-14. (PDF)

Clyde, W. C.  2001.  Mammalian biostratigraphy of the McCullough Peaks area in the northern Bighorn Basin. In Gingerich P. D. (ed.), Paleocene-Eocene Stratigraphy and Biotic Change in the Bighorn and Clarks Fork Basins, Wyoming, University of Michigan Papers on Paleontology 33: 109-126. (PDF)

Gingerich, P. D., Arif, M., Khan, I. H., Haq, M.-U., Bloch, J. I., Clyde, W. C., and Gunnell, G. F.  2001.  Gandhera quarry, a unique mammalian faunal assemblage from the early Eocene of Baluchistan (Pakistan).  In Gunnell, G. F., ed., Eocene Vertebrates: Unusual Occurrences and Rarely Sampled Habitats. Plenum Press. (PDF)

MacLeod, N., Ortiz, N., Fefferman, N., Clyde, W., Schulter, C. and MacLean, J.  2000.  Phenotypic response of foraminifera to episodes of global environmental change. In Culver, S. J. and Rawson, P. (eds.),  Biotic Response to Global Change, Cambridge University Press.

Gingerich, P. D., Arif, M., Bhatti, M. A., and Clyde, W. C.  1998.  Marine mammals (Cetacea and Sirenia) from the middle Eocene of the Sulaiman Range, Pakistan. In Beard, K. C. and Dawson, M. (eds.), Dawn of the Age of Mammals in Asia, Bulletin of the Carnegie Museum of Natural History 34:239-259.

Workshop Proceedings

Walton, A. W., Miller, K. G., Koeberl, C., Shervais, J., Colman, S., Hickman, S., Clyde, W., 2009, The Future of Continental Scientific Drilling: U.S. Perspective; Proceedings of a DOSECC Workshop.